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23.7:

๋ฐฐ์„ค๊ณ„ ๋น„๊ต

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– [๊ฐ•์‚ฌ] ์ƒ๋ฌผํ•™์  ๊ณ ๋ถ„์ž์ธํƒ„์ˆ˜ํ™”๋ฌผ, ์ง€์งˆ, ๋‹จ๋ฐฑ์งˆ ๋ฐ ํ•ต์‚ฐ์€๋ชธ์—์„œ ๊ฐ€์žฅ ์ค‘์š”ํ•œ ๋Œ€ํ˜• ๋ถ„์ž์ž…๋‹ˆ๋‹ค๋‹จ๋ฐฑ์งˆ๊ณผ ํ•ต์‚ฐ์— ํ•จ์œ ๋œ ์งˆ์†Œ๋Š”์ข…์ข… ๋ถ€์‚ฐ๋ฌผ์˜ ๊ฒฐ๊ณผ๋กœ์„œ์ด๋Ÿฌํ•œ ๋ถ„์ž๋“ค์˜ ๋ถ„ํ•ด ์‹œ ๋ฐœ์ƒํ•ฉ๋‹ˆ๋‹ค์‹ ์ฒด์˜ ๊ณผ๋„ํ•œ ์งˆ์†Œ๋Š” ์•”๋ชจ๋‹ˆ์•„๋ฅผ ํ˜•์„ฑํ•˜๊ฒŒ ๋˜๋ฉฐ์ด๋Š” ๋…์„ฑ์ด ๊ฐ•ํ•˜๋ฉฐ ์ œ๊ฑฐ ๋ฐฉ์‹์€ ์ง์ ‘์ ์ธ, ๋˜๋Š”์š”์†Œ, ํ˜น์€ ์š”์‚ฐ์œผ๋กœ ์ „ํ™˜๋œ ํ›„ ์‹คํ–‰๋ฉ๋‹ˆ๋‹ค๋Œ€๋ถ€๋ถ„์˜ ์ˆ˜์ƒ ๋™๋ฌผ์€ ์•”๋ชจ๋‹ˆ์•„๋ฅผ์ง์ ‘ ํ•ด๋‹น ํ™˜๊ฒฝ์œผ๋กœ ๋ฐฐ์ถœํ•ฉ๋‹ˆ๋‹ค๋Œ€๋ถ€๋ถ„์˜ ์•”๋ชจ๋‹ˆ์•„๋Š” ํ™•์‚ฐ์œผ๋กœ ์†์‹ค๋˜๋ฉฐ์—๋„ˆ์ง€ ์ง‘์•ฝ์ ๊ณผ๋Š” ๊ฑฐ๋ฆฌ๊ฐ€ ๋จผ ๊ณผ์ •์ด๋‚˜์•”๋ชจ๋‹ˆ์•„๋ฅผ ๊ฒฌ๋”œ ์ˆ˜ ์žˆ๋Š” ์ •๋„๋Š”๋†๋„๊ฐ€ ๋‚ฎ์€ ๊ฒฝ์šฐ์ด๋ฉฐ์ด์— ๋Œ€ํ•œ ํฌ์„ ๊ณผ์ •์„ ์œ„ํ•ด ํ•ด๋‹น ๋™๋ฌผ์—๊ฒŒ ๋งŽ์€ ์–‘์˜ ๋ฌผ์ด ์š”๊ตฌ๋ฉ๋‹ˆ๋‹ค๋งŽ์€ ์ƒ๋ฌผ์ฒด์˜ ๊ฒฝ์šฐ, ๋ฌผ์˜ ์†Œ๋น„๋Š” ๋งค์šฐ ์ƒ๋‹นํ•ฉ๋‹ˆ๋‹คํฌ์œ ๋ฅ˜, ๋งŽ์€ ์„ฑ์ฒด ์–‘์„œ๋ฅ˜ ๋ฐ์ผ๋ถ€ ํ•ด์–‘ ์ƒ๋ฌผ์€ ์•”๋ชจ๋‹ˆ์•„๋ฅผ ์š”์†Œ๋กœ ์ „ํ™˜ํ•˜๋Š”๋ฐ๋ชธ์—์„œ ๋ฐฉ์ถœ๋˜๊ธฐ ์ด์ „์— ์‹คํ–‰ํ•ฉ๋‹ˆ๋‹ค์š”์†Œ๋Š” ์•”๋ชจ๋‹ˆ์•„๋ณด๋‹ค ๋…์„ฑ์ด ํ›จ์”ฌ ์ ์œผ๋ฉฐ์ ์€ ์–‘์˜ ๋ฌผ์ด ์ œ๊ฑฐ์— ํ•„์š”ํ•ฉ๋‹ˆ๋‹ค๊ทธ๋Ÿฌ๋‚˜, ์•”๋ชจ๋‹ˆ์•„์˜ ์š”์†Œ ์ „ํ™˜์€ ์—๋„ˆ์ง€๊ฐ€ ์š”๊ตฌ๋ฉ๋‹ˆ๋‹ค์กฐ๋ฅ˜, ํŒŒ์ถฉ๋ฅ˜ ๋ฐ ๊ณค์ถฉ์— ์˜ํ•ด ์•”๋ชจ๋‹ˆ์•„๋Š”์ฃผ๋กœ ์š”์‚ฐ์œผ๋กœ ์ „ํ™˜๋ฉ๋‹ˆ๋‹ค์š”์‚ฐ์€ ๋” ๋‹จ๋‹จํ•œ ํ˜•ํƒœ๋กœ ๋ฐฐ์„ค ๋  ์ˆ˜ ์žˆ์œผ๋ฉฐ๋”ฐ๋ผ์„œ ๋ฌผ์œผ ํ•„์š”์„ฑ์€ ๊ฑฐ์˜ ์—†์œผ๋‚˜์•”๋ชจ๋‹ˆ์•„์˜ ์š”์‚ฐ ์ „ํ™˜ ๊ณผ์ •์€์š”์†Œ์˜ ๊ฒฝ์šฐ๋ณด๋‹ค ํ›จ์”ฌ ๋” ์—๋„ˆ์ง€ ์ง‘์•ฝ์ ์ธ ๊ณผ์ •์ž…๋‹ˆ๋‹ค์ด๋Ÿฌํ•œ ์•”๋ชจ๋‹ˆ์•„ ์ œ๊ฑฐ ๋ฐฉ๋ฒ•์˜ ์žฅ์ ๊ณผ ๋Œ€๊ฐ€๋Š”์ฆ‰, ์š”์†Œ ๋˜๋Š” ์š”์‚ฐ์œผ๋กœ์˜ ์ง์ ‘ ๋ฐฉ์ถœ ๋˜๋Š” ์ „ํ™˜์€๋‹ค๋ฅธ ์„œ์‹์ง€ ํ™˜๊ฒฝ์— ๋Œ€ํ•œ ์œ ๊ธฐ์ฒด์˜ ์ ์‘์„ ๋ฐ˜์˜ํ•ฉ๋‹ˆ๋‹คํ™”์„ ์ฆ๊ฑฐ๋Š” ์ƒ๋ช…์ด ๋ฌผ์—์„œ ์‹œ์ž‘๋˜์—ˆ์Œ์„ ๋ณด์—ฌ์ค๋‹ˆ๋‹ค์œ ๊ธฐ์ฒด๊ฐ€ ์œก์ง€๋กœ ์ด๋™ํ•จ์— ๋”ฐ๋ผ ๊ฑด์กฐํ•œ ์กฐ๊ฑด์œผ๋กœ ์ธํ•ด์š”์‚ฐ ๊ฒฝ๋กœ์˜ ์ง„ํ™”๋กœ ์ด์–ด์ ธ๋” ๋งŽ์€ ๋ฌผ์ด ๋™๋ฌผ์— ์˜ํ•ด ๋ณด์กด๋  ์ˆ˜ ์žˆ์—ˆ์Šต๋‹ˆ๋‹ค์ƒ์ดํ•œ ์ƒ์‹์  ํŠน์„ฑ ๋˜ํ•œ์งˆ์†Œ์„ฑ ๋…ธํ๋ฌผ ์ œ๊ฑฐ์— ์žˆ์–ด ๊ณ ์œ ์˜ ๋ฐฉ๋ฒ•์„ ํƒํ•˜๊ฒŒ ํ–ˆ์œผ๋ฉฐ์˜ˆ๋ฅผ ๋“ค์–ด, ์š”์†Œ์˜ ์ˆ˜์šฉ์„ฑ์œผ๋กœ ์ธํ•˜์—ฌํฌ์œ ๋ฅ˜ ๋ฐฐ์•„๊ฐ€ ๋…ธํ๋ฌผ์„๋ชจ์ฒด์˜ ํ˜ˆ์•ก์— ์ œ๊ฑฐํ•  ์ˆ˜ ์žˆ์Šต๋‹ˆ๋‹คํ•œํŽธ, ์š”์‚ฐ์˜ ๊ฒฌ๊ณ ํ•œ ์„ฑ์งˆ์€์กฐ๋ฅ˜ ๋ฐฐ์•„์˜ ๋…ธํŽ˜๋ฌผ์€์•Œ ๋‚ด๋ถ€์˜ ๋ฌดํ•ดํ•œ ๋ฉ์–ด๋ฆฌ ํ˜•ํƒœ๋ฅผ ๊ฐ€๋Šฅ์ผ€ ํ•˜๋ฉฐ๋”ฑ๋”ฑํ•œ ๊ป์งˆ ๋•Œ๋ฌธ์—์š”์†Œ๋Š” ํˆฌ๊ณผ๋  ์ˆ˜ ์—†์Šต๋‹ˆ๋‹ค

23.7:

๋ฐฐ์„ค๊ณ„ ๋น„๊ต

๋™๋ฌผ๋“ค์€ ๋ฐฐ์„ค, ์ฆ‰ ๋ชธ ์†์˜ ๋…ธํ๋ฌผ ์ œ๊ฑฐ์— ๋Œ€ํ•œ ๋‹ค์–‘ํ•œ ์ „๋žต์„ ์ง„ํ™” ์‹œ์ผœ ์™”์Šต๋‹ˆ๋‹ค. ๋Œ€๋ถ€๋ถ„์˜ ๋…ธํ๋ฌผ์€ ๋ฐฐ์„ค๋˜๊ธฐ ์œ„ํ•ด ๋ฌผ์— ์šฉํ•ด๋˜์–ด์•ผ ํ•˜๋ฏ€๋กœ ๋™๋ฌผ์˜ ๋ฐฐ์„ค ์ „๋žต์€ ๋ฌผ ๊ท ํ˜•์— ์ง์ ‘์ ์ธ ์˜ํ–ฅ์„ ๋ฏธ์นฉ๋‹ˆ๋‹ค.

์งˆ์†Œ์„ฑ ๋…ธํ๋ฌผ์€ ๋™๋ฌผ ๋…ธํ๋ฌผ์˜ ๊ฐ€์žฅ ์ค‘์š”ํ•œ ํ˜•ํƒœ ์ค‘ ํ•˜๋‚˜์ž…๋‹ˆ๋‹ค. ์งˆ์†Œ๋Š” ๋‹จ๋ฐฑ์งˆ๊ณผ ํ•ต์‚ฐ์„ ์—๋„ˆ์ง€ ๋˜๋Š” ํƒ„์ˆ˜ํ™”๋ฌผ๊ณผ ์ง€๋ฐฉ์œผ๋กœ ์ „ํ™˜ํ•˜๊ธฐ ์œ„ํ•ด ๋ถ„ํ•ดํ•  ๋•Œ ๋ฐฉ์ถœ๋ฉ๋‹ˆ๋‹ค. ๋‹จ๋ฐฑ์งˆ์€ ์•„๋ฏธ๋…ธ์‚ฐ์œผ๋กœ, ํ•ต์‚ฐ์€ ์งˆ์†Œ ์—ผ๊ธฐ๋กœ ๋ถ„ํ•ด๋ฉ๋‹ˆ๋‹ค. ์•„๋ฏธ๋…ธ์‚ฐ๊ณผ ์งˆ์†Œ ์—ผ๊ธฐ์—์„œ ์งˆ์†Œ๋ฅผ ํฌํ•จํ•˜๋Š” ์•„๋ฏธ๋…ธ๊ธฐ๋Š” ์งˆ์†Œ์„ฑ ๋…ธํ๋ฌผ๋กœ ๋ณ€ํ™˜๋ฉ๋‹ˆ๋‹ค.

๋™๋ฌผ์ด ๋ฐฐ์ถœํ•˜๋Š” ์ „ํ˜•์ ์ธ ์งˆ์†Œ์„ฑ ๋…ธํ๋ฌผ์€ ์•”๋ชจ๋‹ˆ์•„, ์š”์†Œ(urea), ์š”์‚ฐ(uric acid)์ž…๋‹ˆ๋‹ค. ์ด๋Ÿฌํ•œ ๋ฐฐ์„ค ์ „๋žต์€ ์—๋„ˆ์ง€ ์ ˆ์•ฝ๊ณผ ๋ฌผ ์ ˆ์•ฝ ์‚ฌ์ด์˜ ์ ˆ์ถฉ์„ ๋ณด์—ฌ์ค๋‹ˆ๋‹ค.

๋‹ค์–‘ํ•œ ์งˆ์†Œ์„ฑ ๋…ธํ๋ฌผ์€ ๋šœ๋ ทํ•œ ์„œ์‹์ง€์™€ ์ง„ํ™” ์—ญ์‚ฌ๋ฅผ ๋ฐ˜์˜ํ•ฉ๋‹ˆ๋‹ค. ์˜ˆ๋ฅผ ๋“ค์–ด, ๋Œ€๋ถ€๋ถ„์˜ ์ˆ˜์ƒ ๋™๋ฌผ์€ ์•”๋ชจ๋‹ˆ์•„๋ฅผ ์ง์ ‘ ๋ฐฐ์„คํ•ฉ๋‹ˆ๋‹ค (ammonotelic). ์ด ์ ‘๊ทผ ๋ฐฉ์‹์€ ์•”๋ชจ๋‹ˆ์•„๋ฅผ ๋ฐฐ์„คํ•˜๊ธฐ ์ „์— ์š”์†Œ๋‚˜ ์š”์‚ฐ์œผ๋กœ ๋ณ€ํ™˜ํ•˜๋Š” ๊ฒƒ๋ณด๋‹ค ์—๋„ˆ์ง€๋ฅผ ๋œ ์†Œ๋ชจํ•˜์ง€๋งŒ, ๋” ๋งŽ์€ ๋ฌผ์ด ํ•„์š”ํ•ฉ๋‹ˆ๋‹ค. ์œก์ง€ ์ƒ๋ฌผ์—๊ฒ ํƒˆ์ˆ˜๋ณด๋‹ค ๋” ์œ„์ค‘ํ•œ ์œ„ํ˜‘์€ ๋“œ๋ฌผ๊ธฐ ๋•Œ๋ฌธ์— ๋ฌผ ๋ณด์กด์„ ์œ„ํ•ด ์ถ”๊ฐ€์ ์ธ ์—๋„ˆ์ง€ ๋น„์šฉ์„ ๋‚ผ ๊ฐ€์น˜๊ฐ€ ์žˆ์Šต๋‹ˆ๋‹ค.

ํฌ์œ ๋ฅ˜๋‚˜ ์ƒ์–ด ๊ฐ™์€ ๋™๋ฌผ์€ ๋ฐฐ์„คํ•˜๊ธฐ ์ „์— ์•”๋ชจ๋‹ˆ์•„๋ฅผ ์š”์†Œ๋กœ ๋ณ€ํ™˜ํ•ฉ๋‹ˆ๋‹ค (ureotelic). ์š”์†Œ๋Š” ์•”๋ชจ๋‹ˆ์•„๋ณด๋‹ค ๋…์„ฑ์ด ์ ๊ณ  ์ œ๊ฑฐ๋ฅผ ์œ„ํ•ด ์“ฐ์ด๋Š” ๋ฌผ์ด ๋œ ํ•„์š”ํ•ฉ๋‹ˆ๋‹ค. ์ˆ˜์ƒ ์„œ์‹์ง€์—์„œ ์œก์ง€ ์„œ์‹์ง€๋กœ ์ด๋™ํ•˜๋Š” ๋งŽ์€ ์–‘์„œ๋ฅ˜๋Š” ์˜ฌ์ฑ™์ด์ผ ๋• ์ฃผ๋กœ ์•”๋ชจ๋‹ˆ์•„๋ฅผ ๋ฐฐ์„คํ•˜์ง€๋งŒ, ์œก์ง€์˜ ์„ฑ์ธ์ผ ๋• ๋Œ€๋ถ€๋ถ„ ์š”์†Œ๋ฅผ ๋ฐฐ์„คํ•ฉ๋‹ˆ๋‹ค.

ํŒŒ์ถฉ๋ฅ˜, ์ƒˆ, ๊ทธ๋ฆฌ๊ณ  ๋งŽ์€ ๊ณค์ถฉ์„ ํฌํ•จํ•œ ์œ ๊ธฐ์ฒด๋Š” ๋ฐฐ์„คํ•˜๊ธฐ ์ „์— ์•”๋ชจ๋‹ˆ์•„๋ฅผ ์š”์‚ฐ์œผ๋กœ ๋ณ€ํ™˜ํ•ฉ๋‹ˆ๋‹ค (uricotelic). ์š”์‚ฐ์€ ์ˆ˜์šฉ์„ฑ์ด ์—†์œผ๋ฉฐ ๋”ฐ๋ผ์„œ ๋ฌผ์„ ๊ฑฐ์˜ ์‚ฌ์šฉํ•˜์ง€ ์•Š๋Š” ๋ฐ˜์ฃฝ์ด๋‚˜ ๊ฐ€๋ฃจ๋กœ ๋ฐฐ์„ค๋ฉ๋‹ˆ๋‹ค. ์š”์‚ฐ์€ ์š”์†Œ๋ณด๋‹ค ๋…์„ฑ์ด ๋œํ•ฉ๋‹ˆ๋‹ค. ํ•˜์ง€๋งŒ ์•”๋ชจ๋‹ˆ์•„๋ฅผ ์š”์‚ฐ์œผ๋กœ ๋ณ€ํ™˜ํ•˜๋Š” ๊ฒƒ์€ ์š”์†Œ๋กœ ๋ณ€ํ™˜ํ•˜๋Š” ๊ฒƒ๋ณด๋‹ค ํ›จ์”ฌ ๋” ๋งŽ์€ ์—๋„ˆ์ง€๊ฐ€ ํ•„์š”ํ•ฉ๋‹ˆ๋‹ค.

์œ„์™€ ๊ฐ™์€ ๋‹ค์–‘ํ•œ ๋ฐฐ์„ค ์ „๋žต์€ ๋™๋ฌผ๋“ค์ด ์ž์‹ ์˜ ํ™˜๊ฒฝ์—์„œ ๋ฌผ๊ณผ ์—๋„ˆ์ง€ ์ˆ˜์š”๋ฅผ ์ถฉ์กฑ์‹œํ‚ฌ ์ˆ˜ ์žˆ๊ฒŒ ํ•ด์ค๋‹ˆ๋‹ค.

Suggested Reading

Wright, P A. “Nitrogen Excretion: Three End Products, Many Physiological Roles.” The Journal of Experimental Biology 198, no. 2 (February 1, 1995): 273.[Source]>

Spring, Jeffrey H., S. Renee Robichaux, and John A. Hamlin. “The Role of Aquaporins in Excretion in Insects.” Journal of Experimental Biology 212, no. 3 (February 1, 2009): 358. [Source]